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Scientific Name:
Styela clavaCommon Name:
Club tunicate (also: clubbed tunicate, leathery sea squirt, Asian sea squirt)Native Range:
JapanEstablished Range:
S. clava has established itself off the coasts of Australia, New Zealand, Europe, and North America. On the East Coast, it is found from Canada to Maryland.Established in Rhode Island?
Yes, throughout Narragansett BayDate and Location of Introduction:
1970, Beverly, MassachusettsMethod of Introduction:
Unknown, but likely ballast water or hull foulingHabitat:
S. clava generally lives in the subtidal zone, attached to rocks, pilings, floating docks, and the hulls of ships. It can tolerate water temperatures from -2 to 23° Celcius, and salinities between 22 and 36 ppt.Average Life Span:
1 to 3 yearsDiet:
S. clava is a filter feeder that eats zooplankton, phytoplankton and detritus.Breeding:
S. clavais hermaphroditic and breeds by broadcast spawning. In New England, it breeds between June and November.Concerns:
All invasive tunicates, including S. clava, pose the same problems. These tunicates are notorious fouling organisms, and can completely cover submerged boat hulls, aquaculture cages, and just about any other surface that they are capable of living on. As a result, they can slow down boats and have negative impacts on the local environment. Invasive tunicates have been known to smother shellfish and other sessile organisms, and will outcompete native filter feeders for food and space.Control:
S. clavacan be removed from an infested object by scraping it off
by hand and placing it in the garbage or letting it dry out. If you choose to
pressure wash it off of equipment, only do so on land and make sure the
resulting wastewater does not go back into the sea. Additionally, S.
clavacan be removed from smaller objects, such as shellfish, by placing it
in vinegar or a brine bath and then exposing it to air.
Identification Card:
Documents:
- WASHINGTON STATE’S RESPONSE TO AN INVASION OF NON-NATIVE TUNICATES
- Biological Synopsis of the Invasive Tunicate Styela clava (Herdman 1881)
Works Cited:
Cohen, Andrew N. 2005. Guide to the Exotic Species of San Francisco Bay. San Francisco Estuary Institute, Oakland, CA, www.exoticsguide.orgFuller, Pam. 2010. Styela clava. USGS Nonindigenous Aquatic Species Database, Gainesville, FL. http://nas.er.usgs.gov/queries/FactSheet.aspx?speciesID=1292
Clarke, C.L., and T.W. Therriault. Biological Synopsis of the Invasive Tunicate Styela clava(Herdman 1881). Fisheries and Oceans Canada, Nanaimo, BC. http://www.dfo-mpo.gc.ca/Library/329953.pdf

